Transformers: Devastation is an action game based on the Transformers franchise, specifically the original cartoon series from the 1980s now known as Generation 1 . It has the same bright visual style using cel-shaded graphics. The main story mode has five difficulty levels and tells how the Decepticons plan to cyberform planet Earth after Megatron finds a way to control plasma energy. He uses this technique to turn organic material into metal. The Autobots need to prevent this and eventually five different characters are playable: Bumblebee, Grimlock, Optimus Prime, Sideswipe and Wheeljack.
The gameplay is similar to other titles by developer Platinum Games. The game is largely played as a 3D action brawler where missions are divided into smaller areas with short cut-scenes in between to further the story. The Autobots link light and heavy attack in different patterns to form combos. When a blue flash appears during a combo it can be finished off with an attack in vehicle form. Robots can freely switch between robot and vehicle form.
The game features a variety of weapons, melee and ranged, which can be collected during missions and switched on the fly. Ranged weapons have an energy meter and when empty require some time to recharge. There are also transpatial converters that temporary reveal hidden items in another location. The Autobots can also pick up and throw objects to solve small environment puzzles.
Encounters with large robot bosses feature weak points that need to be exploited, and the game has a ranking system based on performance after each fight. Health and weapon energy can be restored using green and blue energon cubes. Players can select their Autobot and define their loadout from a large inventory of weapons.
The credits currency earned during missions can be given to Wheeljack, who develops T.E.C.H. that provides enhanced abilities. These enhancements require quick timing mini-games to develop and can be selected for two slots. Equipped T.E.C.H. can be different for each mission.