The player takes on the role of Dee Dee, a skilled female pilot who starts out smuggling whiskey to Allied forces but eventually joins the fight against Axis armies. The campaign spans from 1941 to 1945, taking her to Africa and Europe. While the story primarily focuses on World War II, some flashback missions also take place during World War I. During WWII missions, the player earns points by destroying enemy targets, which can be used to buy upgrades such as plane endurance or leadership.
By completing campaign missions, new planes are unlocked, including British, German, and Russian aircraft. Missions vary in nature and consist of escorting bombers, protecting ground troops, dogfights, and more. The player's main weapons are machine guns, bombs, and rockets, with some limitations on each type depending on the plane. Ground targets are generally easier to hit than enemy planes, but luck plays a role when using rockets. The bomb or rocket camera can be activated by holding the fire button to see if the target has been hit.
The game features both behind-the-plane and 1st-person views, with options for different types of combat, locations, and weather conditions. This allows for flexibility in playing through the game's various missions.