Transformers: Devastation is an action game based on the Transformers franchise, specifically the original cartoon series from the 1980s now known as Generation 1 . It has the same bright visual style using cel-shaded graphics. The main story mode has five difficulty levels and tells how the Decepticons plan to cyberform planet Earth after Megatron finds a way to control plasma energy. He uses this technique to turn organic material into metal. The Autobots need to prevent this and eventually five different characters are playable: Bumblebee, Grimlock, Optimus Prime, Sideswipe and Wheeljack.
The gameplay is similar to other titles by developer Platinum Games. The game is largely played as a 3D action brawler where missions are divided into smaller areas with short cut-scenes in between to further the story. The Autobots link light and heavy attack in different patterns to form combos. When a blue flash appears during a combo it can be finished off with an attack in vehicle form. Robots can freely switch between robot and vehicle form.
The gameplay also features longer driving sequences, often when chasing or escaping. While driving, ranged weapons can be fired and there is a rush attack and obstacles to dodge and booster portals to receive extra speed. Melee weapons include swords, gauntlets and hammers and ranged weapons are blasters, grenade launchers etc.
During gameplay green and blue energon cubes can be picked up to restore health and weapon energy respectively. New missions are always started at the Ark headquarters. The player gets to select the Autobot and can define the loadout from the large inventory of weapons. Weapons have their own levels and can be made stronger through synthesis, merging weapons with a base one and the secondary ones used for parts.
The game has many encounters with large robot bosses where weak points need to be exploited. After each fight a ranking is shown based on the performance. Players also get to develop new abilities by completing quick timing mini-games, which can then be selected from a large pool for two slots.