Players take on the role of leading a small force of mechs in a fight to save humanity from an alien invasion in Super Robot Taisen Original Generation . The game's story picks up six months after the events of its predecessor, with most people on Earth still trying to recover from their battle against the Aerogaters. A small faction of the Divine Crusaders remains determined to further their goals.
The gameplay is turn-based, with players controlling multiple mechs and issuing commands in a similar style to its predecessor. The combat system has been improved, with new attack types such as chain attacks allowing units to strike multiple enemies in one blow. The AI has also been enhanced, making battles more challenging and dynamic.
Meach pilots can also use "spirit commands" to strengthen their mechs' abilities, such as increasing its hit or dodge rate. Players must carefully manage their mechs' movements and attacks to outmaneuver their enemies. The game's graphics have been enhanced, with smoother motion and more detailed attacks from the mechs.